College of Agronomy and Biotechnology, Yunnan Agricultural University, 650201 Kunming, China. Institute of Medicinal Plants, Yunnan Academy of Agricultural Sciences, 650200 Kunming, China. College of Resources and Environment, Yuxi Normal University, 653100 Yuxi, China. Institute of Medicinal Plants, Yunnan Academy of Agricultural Sciences, 650200 Kunming, China. Electronic address: boletus@126.com. College of Agronomy and Biotechnology, Yunnan Agricultural University, 650201 Kunming, China. Electronic address: honggaoliu@126.com.